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Epitoniidae Berry, 1910 (1812)

Cirsotrema varicosum

(Lamarck, 1822)

Description:

Dimensions range from 9 mm - 70.2 mm high and 5.2 mm - 33 mm wide.
Width to height ratio: 0.58:1 ~ 0.47:1

Shell: slender conical, turreted; thick, very robust; small to medium. Exterior colour blueish white, white or sand coloured. About 12 whorls, convex, compressed, distinct; protoconch creamy white or yellowish, smooth; teleoconch with 12-11 whorls more or less prominent, the last one encircled by an anterior keel. Spire tall, acute. Suture deep, channelled, interrupted by extending laminae. Imperforate: umbilicus absent. Approximately 16 - 21 costae on the body whorl. Costae are small, flattened, wavelike and finely crenulate, the roughness of the fringes are sharper towards the suture, prominent at the lower keel. There are 1 - 2 varices arranged randomly on the shell. They are large, thick, crenulated, roughly aligned up the side, and bear thick rugae where the spiral ribs cross. Costae not in line on adjoining whorls; often coronate, raised and produced at the suture; few peaks merged with costae on preceding whorl. Intercostal spaces spiral ribs form a cancellate pattern where they cross the costae with small, but deep pits in the interstices. Base with thick spiral ridge consisting of overlapping, flattened nodules situated close to the columella. Columella thick, with a deep slot between columella and rib. Aperture rounded, with a thick margin; inner lip expanded and reflected.


Distribution:

Hawaii and Polynesia to northern Mozambique [Kilburn, 1985:255]; Mauritius; Indonesia [Kokshoorn et al. 2007: 36]; Off Kii Peninsula to Okinawa, Japan [Nakayama, 2003:23-24]; Abrolhols, WA to Southern QLD [Wilson, 1993:274], NSW, Tasmania, New Caledonia [Dushane, 1990:3]


Habitat:

intertidal to 270m on sand & rubble substrate underneath the anemones Stichodactyla haddoni, Actinodendron plumosum, Actinodendron arboreum, Heteractis crispa and Macrodactyla doreensis


Etymology:

[latin] from varicosus = varicose, full of dilated veins, possibly in reference to the appearance of the sculpture


Type Material:

Syntype: BMNH 198144/1 [as Cirsotrema multiperforatum (G. B. Sowerby II, 1874)] - Type Locality: Black River Bay, Mauritius. [Kokshoorn, B., Gittenberger, J. & Gittenberger, A, 2007]
Type not located [as Scalaria abbreviata G.B. Sowerby II, 1874: ~ Scala (Cirsotrema) joubini de Boury, 1913c.] - Type Locality: Tamarind Bay, near Black River, Mauritius.. [Brown, L. & Neville, B.D., 2015]
Holotype: MNHN 4222 [as Scala bavayi de Boury, 1913:] - Type Locality: New Caledonia.. [Brown, L. & Neville, B.D., 2015]
Type not located [as Scalaria fimbriata Lamarck, 1816:] - Type Locality: none given. [Brown, L. & Neville, B.D., 2015]
Holotype: NHMUK 1902.11.26.71 [as Scalaria multiperforata G.B. Sowerby II, 1874:] - Type Locality: Black River Bay, Mauritius.. [Brown, L. & Neville, B.D., 2015]
Holotype: MNHN-IM-2000-4222 Muséum national d'Histoire naturelle [as Scala bavayi de Boury, 1912] - Type Locality: New Caledonia. [MNHN, 2025c]
Type not listed in source data for Cirsotrema varicosum (Lamarck, 1822) - Type Locality: none designated. [Higo, S., Callomon, P. & Goto, Y., 1999]
Type not listed in source data [as Cirsotrema abbreviatum (G. B. Sowerby II, 1874)] - Type Locality: Philippines. [Higo, S., Callomon, P. & Goto, Y., 1999]
Type not listed in source data for Cirsotrema varicosum (Lamarck, 1822) - Type Locality: 'Nouvelle-Calédonie' (New Caledonia).. [Higo, S., Callomon, P. & Goto, Y., 1999]


SpeciesImage

Murrell (2025).

off Broome, Western Australia.
Trawled by scampi boats in muddy sand from 450-480m.

Distribution

as listed in source literature

Sowerby, G. B. II. (1844).
Species Image
Plate xxxv. Fig. 126.
Species Image
Plate xxxv. Fig. 128.
Species Image
Plate xxxv. Fig. 143.
Sowerby, G. B. II. (1873).
Species Image
Plate ii. Fig. 11a.
Species Image
Plate ii. Fig. 11b.
Sowerby, G. B. II. (1873).
Species Image
Plate xiv. Fig. 125.
described as Scalaria multiperforata G. B. Sowerby II, 1874
Sowerby, G. B. II. (1873).
Species Image
Plate xiv. Fig. 124.
described as Scalaria abbreviata G. B. Sowerby II, 1874
Tryon, G.W. (1887).
Shown in text as Scalaria varicosa Lamarck, 1822, Scalaria abbreviata G. B. Sowerby II, 1874
Species Image
Plate 17. Fig. 23.
Species Image
Plate 17. Fig. 27.
Scalaria abbreviata G. B. Sowerby II, 1874
Species Image
pp: 81.
Clessin, S. (1897).
Species Image
Plate 3. Fig. 7.
Species Image
Plate 1. Fig. 2.
The illustration on our Plate 1, Fig. 2 is based on a specimen from the Stuttgart Natural History Museum, in which the longitudinal folds are more clearly developed than in the depiction of the species given by Reeve (Plate 3, Fig. 7).
Species Image
pp: 9-10.
Shell white, thick, ornamented with nodular spiral ridges; whorls distinct, the last keeled spirally below; suture deep, frequently interrupted by projecting lamellae; varices very rare, thick, crenulated; aperture round, outer lip expanded and reflexed above and below.
Clessin, S. (1897).
Species Image
Plate 4. Fig. 6.
Species Image
pp: 16.
Shell short, thick, somewhat bluish, imperforate, spirally grooved; ribs flat, undulating, angled and looped on one side, spined at the suture, and projecting at the lower keel; interstices deeply pitted at the suture and keel; whorls few, somewhat sloping above; last whorl keeled below, thickened at the aperture margin; varices large, thick, crenulated, very few; aperture nearly round, margin thick and crenate.
Kilburn, R.N (1985).
Species Image
pp: 255. Plate Fig 21, 22 .
21. Cirsotrema (Cirsotrema) varicosum (Lamarck, 1822),
NM H4857, Conducia Bay, Mozambique, 41,9x ÍS,8 mm;

22, C.varicosum, NM H4856, Conducia Bay, 34,1 x 14,2 mm.
Nakayama, T. (2003).
Species Image
pp: 109. Plate 3. Fig. 17-20.
figs 17-19: Cirsotrema (Cirsotrema) varicosum (Lamarck, 1822)
height 70.2 mm; breadth 23.0 mm (NA); fig. 19, base
fig. 20:
height 9.0 mm; breadth 5.2 mm (ND)
Kokshoorn, B., Gittenberger, J. & Gittenberger, A (2007).
Species Image
pp: 38. Fig. 2-3.
described as Cirsotrema multiperforata (Sowerby [II],1874)
2 syntype (BMNH198144), H=16.3mm;
3 E.side of Samalona island (RMNH87814),H=18.1mm
Kokshoorn, B., Gittenberger, J. & Gittenberger, A (2007).
Species Image
pp: 38. Plate 4-4a.
4 Indonesia, Bali, NE.side of Pulau Serangan island, 22m depth, RMNH91205, H=15.0mm;
4a, H=26.6 mm;
Species Image
pp: 38. Plate 5a.
C.varicosum (Lamarck,1822)
5a W.side of Kudingareng Keke island (RMNH87813),H=54.7mm;
Species Image
pp: 38. Plate 5-6.
C.varicosum (Lamarck,1822)
5a W.side of Kudingareng Keke island (RMNH87813),H=54.7mm;
6 W.side of Kudingareng Keke island (RMNH 87771),H=47.1 mm).
Murrell (2025).
Species Image
off Broome, Western Australia.
Trawled by scampi boats in muddy sand from 450-480m.
MNHN (2025c).
Species Image
Holotype Scala bavayi de Boury, 1912. MNHN-IM-2000-4222

Synonymy:

Cirsotrema (Elegantiscala) varicosum 'Cahn (MS)' Azuma, 1960 [in Brown, L. & Neville, B.D., 2015].
Cirsotrema abbreviatum (G. B. Sowerby II, 1874) [in Higo, S., Callomon, P. & Goto, Y., 1999].
Cirsotrema joubini de Boury, 1911: [in Brown, L. & Neville, B.D., 2015].
Cirsotrema multiperforatum (G. B. Sowerby II, 1874) [in Kokshoorn, B., Gittenberger, J. & Gittenberger, A, 2007].
Scala bavayi de Boury, 1912 [in MNHN, 2025c].
Scala bavayi de Boury, 1913: [in Brown, L. & Neville, B.D., 2015].
Scala joubini de Boury, 1913: [in Brown, L. & Neville, B.D., 2015].
Scalaria abbreviata G. B. Sowerby II, 1874 [in Clessin, S., 1897].
Scalaria abbreviata G. B. Sowerby II, 1874 [in Sowerby, G. B. II., 1873].
Scalaria abbreviata G.B. Sowerby II, 1874: ~ Scala (Cirsotrema) joubini de Boury, 1913c. [in Brown, L. & Neville, B.D., 2015].
Scalaria fimbriata Lamarck, 1816 [unaccepted] [in Lamarck, J. B. P. A. de M. de, 1816].
Scalaria fimbriata Lamarck, 1816: [in Brown, L. & Neville, B.D., 2015].
Scalaria multiperforata G. B. Sowerby II, 1874 [in Sowerby, G. B. II., 1873].
Scalaria multiperforata G.B. Sowerby II, 1874: [in Brown, L. & Neville, B.D., 2015].
Scalaria varicosa [in Sowerby, G. B. II., 1844].
Scalaria varicosa Lamarck, 1822, Scalaria abbreviata G. B. Sowerby II, 1874 [in Tryon, G.W., 1887].
Turbo fimbriatus, "W. Wood", 1818 [in Coan, E. & Lutaenko, K. & ZHANG, J. & Sun, Q., 2015].
Turbo varicosus "W. Wood", 1828 [in Coan, E. & Lutaenko, K. & ZHANG, J. & Sun, Q., 2015].


Source Literature:

Brown, L. & Neville, B.D. (2015). Catalog of the recent taxa of the families Epitoniidae and Nystiellidae (Mollusca: Gastropoda) with a bibliography of the descriptive and systematic literature. Zootaxa. 3907(1). Magnolia Press Auckland, New Zealand. http://dx.doi.org/10.11646/zootaxa.3907.1.1 [Accessed 22 October 2023]
Clessin, S. (1897). Die Familie der Scalariidae. In W. Kobelt (Ed.), Systematisches Conchylien-Cabinet von Martini und Chemnitz. Neu herausgegeben und vervollständigt. 2(13). Bauer & Raspe, NürnbergBauer & Raspe, Nürnberg. https://www.biodiversitylibrary.org/page/34259905 [Accessed 21 June 2025]
Coan, E. & Lutaenko, K. & ZHANG, J. & Sun, Q. (2015). The molluscan taxa of A. W. Grabau & S. G. King (1928) and their types. Malacologia. 58(1-2). 10.4002/040.058.0206. [Accessed 21 June 2025]
DuShane, H. (1990). Hawaiian Epitoniidae. Hawaiian Shell News. 38(Sup.1). Hawaiian Malacological Society
Higo, S., Callomon, P. & Goto, Y. (1999). Catalogue and Bibliography of the Marine Shell-Bearing Mollusca of Japan.. https://www.researchgate.net/publication/266375656_Catalogue_and_Bibliography_of_the_Marine_Shell-Bearing_Mollusca_of_Japan [Accessed 25 July 2025]
Kaicher, S.D. (1980). Epitoniidae I [Pack 23]. S. D. Kaicher, St. Petersburg, Florida. http://www.femorale.com/kaicher/species.asp?f=Epitoniidae&c=125 [Accessed 8 May 2023]
Kilburn, R.N (1985). The family Epitoniidae (Mollusca: Gastropoda) in southern Africa and Mozambique. Annals of the Natal Museum. 27(1). Natal Museum
Kokshoorn, B., Gittenberger, J. & Gittenberger, A (2007). Epitoniid parasites (Gastropoda, Caenogastropoda, Epitoniidae) and their host sea anemones (Cnidaria,Actiniaria,Ceriantharia) in the Spermonde archipelago, Sulawesi, Indonesia. Basteria. 71(1-3)
Lamarck, [J.-B. M.] de. (1822). Histoire naturelle des animaux sans vertèbres.. Histoire naturelle des animaux sans vertèbres.. 6(2). Lamarck, [J.-B. M.] de.. http://biodiversitylibrary.org/page/13181853 [Accessed 20 March 2025]
Lamarck, J. B. P. A. de M. de (1816). Mollusques et polypes divers.. Tableau encyclopédique et méthodique des trois règnes de la nature,. 23. https://www.biodiversitylibrary.org/page/37162278#page/165/mode/1up [Accessed 13 June 2025]
MNHN (2025c). Holotype Scala bavayi de Boury, 1912. MNHN-IM-2000-4222 . Muséum National d'Histoire Naturelle [MNHN}. http://coldb.mnhnrf/catalognumber/mnhn/im/2000-4222 [Accessed 30 March 2025]
Murrell (2025). Personal Collection
Nakayama, T. (2003). A Review of Northwest Pacific Epitoniids. Monographs of Marine Mollusca. 6. Backhuys Publishers
Sowerby, G. B. II. (1844). Monograph of the genus Scalaria.. Thesaurus conchyliorum, or monographs of genera of shells.. 1 (4). London, privately published.. http://biodiversitylibrary.org/page/11076419 [Accessed 22 June 2023]
Sowerby, G. B. II. (1873). Monograph of the genus Scalaria. Conchologia iconica, or illustrations of the shells of molluscous animals, vol. 19, pls 1-16 and unpaginated text.. 19. London. https://www.biodiversitylibrary.org/page/8217819 [Accessed 24 August 2023]
Tryon, G.W. (1887). Manual of conchology, structural and systematic with illustrations of the species. IX. Academy of Natural Sciences. https://www.biodiversitylibrary.org/page/11026889 [Accessed 20 September 2025]
Weil, A., Brown, L. & Neville, B. (1999). The Wentletrap Book. Evolver
Wilson, B.R. (1993). Australian Marine Shells. Australian Marine Shells. 1. Odessey Publications
WoRMS Editorial Board (2025). World Register of Marine Species. https://www.marinespecies.org [Accessed 8 November 2025]

WoRMS direct page link: Open WoRMS record