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Epitoniidae Berry, 1910 (1812)

Epitonium fasciatum

(G. B. Sowerby II, 1844)

Description:

Dimensions range from 12.6 mm - 25 mm high and 6 mm - 6.14 mm wide.
Width to height ratio: 0.48:1 ~ 0.25:1

Shell: narrowly conical, pyramidal; fragile; small to moderately small. Exterior colour white with a diffused brown band, costae white. About 9 whorls, rounded, sub-angulated at the shoulder, barely touching; teleoconch with 8-9 whorls rounded. Suture deeply fenestrate, but not perforated. Umbilicate: fenestrated, partly covered by outer lip. Approximately 6 - 9 costae on the body whorl. Costae are prosocline, thin, sharp-egded, equidistant. Costae continuous; hooked at the shoulders; distinctly recurved. Intercostal spaces smooth. Aperture nearly ovate; peristome wide, double, somewhat auriculate at the base ; outer lip excavated at the suture, then tapering to a sharp point. Operculum nucleus centred, translucent yellowish-brown.

Reeve [1873: pl. 3, fig. 13a-b] noted the species differs from E.alatum in being narrower, the shape of the varices and margin of the aperture. In each species there is an angle at the varix, but in E.fasciatum it is much more pointed and elevated than in E.alatum, and leans towards the whorl., , May also be confused with E. pyramidale or E. philippinarum, both of which have a closed umbilicus [Nakayama, 2003:49]


Distribution:

Off Kii Peninsula, Amami Islands to Okinawa, Japan; Philippines to Australia [Nakayama, 2003]; Pakistan [Melvill & Standen, 1903a:344]; Red Sea to Natal, South Africa [Kilburn, 1985].


Habitat:

10-50 m., fine sand [Higo, et al., 1999: 180].


Etymology:

[latin] fasciatus = banded or striped, in relation to the brown stripe on the shell.


Type Material:

Brown & Neville [2015:71] list Scalaria austrocaledonica Montrouzier, 1859 as a synonym of Epitonium fasciatum (G. B. Sowerby II, 1844); however, WoRMS [2026] recognises Epitonium austrocaledonicum (Montrouzier, 1859) as a distinct species.

Type not located [as Scalaria fasciata G.B. Sowerby II, 1844] - Type Locality: Catanuan, Luzon, Philippines.. [Brown & Neville, 2015: 71]
Type not indicated in text for Epitonium fasciatum (G. B. Sowerby II, 1844) - Type Locality: 'Catanauan, pr. Tayabas, ins. Luzon' [Luzon, Philippines]; 'Found in sandy mud at eight to ten fathoms' [10-50 m]. [Higo et al., 1999: G1887]


SpeciesImage

Sowerby, G. B. II. (1844). Plate xxxii. Fig. 12, 13.

Distribution

as listed in source literature

Sowerby, G. B. II. (1844).
Shown in text as Scalaria fasciata G. B. Sowerby II, 1844
Species Image
Plate xxxii. Fig. 12, 13.
Species Image
pp: 84.
original description
Translation: The shell is similar to that of Scalaria alata but narrower and with a more elevated and acute angle; the aperture is smaller, with an acuminated margin.
Sowerby, G. B. II. (1873).
Shown in text as Scalaria fasciata G. B. Sowerby II, 1844
Species Image
Plate III. Fig. 13a.
Species Image
pp: unpaginated.
Translation: Shell white, with a single spiral brown band between the varices, pyramidal; whorls rounded, scarcely contiguous; varices equidistant, lamellar, projecting, elevated near the suture into an acute angle; aperture somewhat ovate, outer lip excavated at the suture, then becoming sharply pointed nearby.
Species Image
Tryon, G.W. (1887).
Shown in text as Scalaria fasciata G. B. Sowerby II, 1844
Species Image
Plate 11. Fig. 41.
Species Image
pp: 55.
Clessin, S. (1897).
Shown in text as Scalaria fasciata G. B. Sowerby II, 1844
Species Image
Plate 5. Fig. 7-8.
Species Image
pp: 21.
Translation: Shell white, with a single brown spiral band between the varices, pyramidal in shape; whorls rounded, slightly convex, closely set; varices evenly spaced, lamellar, prominent, raised near the suture into a sharp angle; aperture nearly ovate, the outer lip excavated at the suture, then tapering upward into a sharp point.
Kilburn, R.N (1985).
Shown in text as Epitonium (Lamelliscala) fasciatum (Sowerby, 1844)
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NM J8582
Pemba, Mozambique
14.5 x 7.8 mm
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NM J7245
Cerf Is., Seychelles,
6.7 x 9.3 mm
Nakayama, T. (2003).
Shown in text as Epitonium (Lamelliscala) fasciatum (Sowerby, 1844)
Species Image
pp: 127. Plate 12. Fig. 1-2.
Epitonium (Lamelliscala) fasciatum (Sowerby, 1844)
height 12.6 mm; breadth 6.0 mm

Synonymy:

Epitonium (Lamelliscala) fasciatum (Sowerby, 1844) [Kilburn, 1985: 287; Nakayama, 2003: 49]
Lamelliscala fasciata (Sowerby, 1844) [Higo et al., 1999: G1887]
Scala fasciata G. B. Sowerby II, 1844 [Melvill & Standen, 1903a: 344]
Scalaria fasciata G. B. Sowerby II, 1844 [Sowerby, 1844: 84; Brown & Neville, 2015: 71; Reeve, 1873: pl. 3, fig. 13a-b; Clessin, 1897: 21; Tryon, 1887: 55]


Source Literature:

Brown, L. & Neville, B.D. (2015). Catalog of the recent taxa of the families Epitoniidae and Nystiellidae (Mollusca: Gastropoda) with a bibliography of the descriptive and systematic literature. Zootaxa. 3907(1). Magnolia Press Auckland, New Zealand. http://dx.doi.org/10.11646/zootaxa.3907.1.1 [Accessed 22 October 2023]
Clessin, S. (1897). Die Familie der Scalariidae. In W. Kobelt (Ed.), Systematisches Conchylien-Cabinet von Martini und Chemnitz. Neu herausgegeben und vervollständigt. 2(13). Bauer & Raspe, NürnbergBauer & Raspe, Nürnberg. https://www.biodiversitylibrary.org/page/34259905 [Accessed 21 June 2025]
Higo, S., Callomon, P. & Goto, Y. (1999). Catalogue and Bibliography of the Marine Shell-Bearing Mollusca of Japan.. https://www.researchgate.net/publication/266375656_Catalogue_and_Bibliography_of_the_Marine_Shell-Bearing_Mollusca_of_Japan [Accessed 25 July 2025]
Kaicher, S.D. (1980). Epitoniidae I [Pack 23]. S. D. Kaicher, St. Petersburg, Florida. http://www.femorale.com/kaicher/species.asp?f=Epitoniidae&c=125 [Accessed 8 May 2023]
Kilburn, R.N. (1985). The family Epitoniidae (Mollusca: Gastropoda) in southern Africa and Mozambique. Annals of the Natal Museum. 27(1). Natal Museum
Melvill, J. C. & Standen, R. (1903a). The genus Scala (Klein) Humphrey, as represented in the Persian Gulf, Gulf of Oman, and North Arabian Sea, with descriptions of new species.. Journal of Conchology. 10. Conchological Society of Great Britain and Ireland. https://www.biodiversitylibrary.org/page/31650162 [Accessed 26 February 2025]
Nakayama, T. (2003). A Review of Northwest Pacific Epitoniids. Monographs of Marine Mollusca. 6. Backhuys Publishers
Reeve, L. A. (1873). Monograph of the genus Scalaria. Conchologia iconica, or illustrations of the shells of molluscous animal. 19. London. https://www.biodiversitylibrary.org/page/8217819 [Accessed 24 August 2023]
Sowerby, G. B. II. (1844). Monograph of the genus Scalaria.. Thesaurus conchyliorum, or monographs of genera of shells.. 1 (4). London, privately published.. http://biodiversitylibrary.org/page/11076419 [Accessed 22 June 2023]
Tryon, G.W. (1887). Manual of conchology, structural and systematic with illustrations of the species. IX. Academy of Natural Sciences. https://www.biodiversitylibrary.org/page/11026889 [Accessed 20 September 2025]
Weil, A., Brown, L. & Neville, B. (1999). The Wentletrap Book. Evolver
Wilson, B.R. (1993). Australian Marine Shells. Australian Marine Shells. Vol.1. Odessey Publications
WoRMS Editorial Board (2025). World Register of Marine Species. https://www.marinespecies.org at VLIZ. [Accessed 8 November 2025]

WoRMS direct page link: Open WoRMS record