Epitonium pallasi neglectum

(A. Adams & Reeve, 1850)

Description:

Dimensions range from 7.31 mm - 40 mm high and 5.08 mm - 27.01 mm wide.
Width to height ratio: 0.69:1 ~ 0.68:1

Shell: stout, broadly pyramidal; small to medium. Exterior colour fawn or pale-brown, costae white. About 8 whorls, separated, joined by the costae; teleoconch loosely coiledwith 4-8 whorls increasing rapidly. Suture deep, perforated. Umbilicate: wide and deep, rib extending into the umbilicus. Approximately 6 - 16 costae on the body whorl. Costae are sharp and high. Costae aligned; either bluntly angulate on the anterior or hooked at the suture; continuous across the suture. Intercostal spaces with microscopic spiral striae. Aperture oval, slightly reflexed on the anterior; peristome complete. Operculum oval.

Axial costae of E. pallasi neglectum are fewer in number and rather roundly shaped at the suture, while those of E. pallasi pallasi are hooked and numerous.

Distribution:

Japan, south to the Philippines and Queensland, Australia

Habitat:

50-230m, sand

Etymology:

[latin] from neglectus = having been neglected, ignored, or overlooked

Type Material:

China for Epitonium pallasi neglectum (A. Adams & Reeve, 1850). [Nakayama, T., 2003]
Holotype: NHMUK 1874.12.11.197 [as Scalaria neglecta A. Adams & Reeve, 1850: ~ Scalaria inclyta Melvill, 1885.] - Locality: China Sea.. [Brown, L. & Neville, B.D., 2015]
Holotype: NMW.1955.158.00078 National Museum Wales [as Scalaria inclyta Melvill, 1885] - Locality: unknown, but probably China Seas. [NMW National Museum Wales, n.d.]
Type not indicated in text for Epitonium pallasi neglectum (A. Adams & Reeve, 1850) - Locality: China Sea. [Higo, S., Callomon, P. & Goto, Y., 1999]

Adams, A. & Reeve, L. A. (1850). Plate VI. Fig. 5.

Distribution

as listed in source literature

Adams, A. & Reeve, L. A. (1850).

Plate VI. Fig. 5.

pp: 51.

original description: Shell conical-turreted, deeply umbilicate, with seven to eight whorls, rounded, smooth, or, under magnification, minutely spirally impressed-striate; with somewhat distant, narrow, elevated, annular ribs, spinose-hooked near the sutures; aperture rounded, lip expanded; flesh-brownish in colour, with white ribs

Sowerby, G. B. II. (1873-1874).

Shown in text as Scalaria neglecta A. Adams & Reeve, 1850

Plate i. Fig. 1.

Fig. sp1.

Tryon, G.W. (1879).

Shown in text as Scalaria neglecta A. Adams & Reeve, 1850

Plate 11. Fig. 34.

Plate 11. Fig. 35.

pp: 55.

Tryon, G.W. (1879).

Shown in text as Scalaria inclyta Melvill, 1885

Plate 11. Fig. 35.

pp: 55.

Clessin, S. (1897).

Shown in text as Scalaria neglecta A. Adams & Reeve, 1850

Plate 7. Fig. 6.

pp: 27.

Shell broadly pyramidal, perforate, pale yellow between the varices; whorls increasing rapidly, disjointed; varices somewhat numerous, rather thin, continuous; near the suture drawn out into spines.

Kaicher, S.D. (1980).

Card #: EP1-2388.

Considered by some workers merely a variety (or synonym) of pallasi and, by others, a good species; it differs by having more and strongly hooked costae per whorl; to 40

Card #: EP1-2388

Wilson, B.R. (1993).

Shown in text as Epitonium neglectum (A. Adams & Reeve, 1850) ·

pp: 379. Fig. 31.

Nakayama, T. (2003).

pp: 123. Plate 10. Fig. 23.

h: 28.8 mm w: 15.0 mm

NMW National Museum Wales (n.d.).

Shown in text as Scalaria inclyta Melvill, 1885

Synonymy:

Epitonium neglectum (A. Adams & Reeve, 1850) · [in Wilson, B.R., 1993].
Scalaria inclyta Melvill, 1885 [in NMW National Museum Wales, n.d.].
Scalaria inclyta Melvill, 1885 [in Tryon, G.W., 1887].
Scalaria inclyta Melvill, 1885: [in Brown, L. & Neville, B.D., 2015].
Scalaria neglecta A. Adams & Reeve, 1850 [in Clessin, S., 1897].
Scalaria neglecta A. Adams & Reeve, 1850 [in Sowerby, G. B. II., 1873-1874].
Scalaria neglecta A. Adams & Reeve, 1850 [in Tryon, G.W., 1887].
Scalaria neglecta A. Adams & Reeve, 1850: ~ Scalaria inclyta Melvill, 1885. [in Brown, L. & Neville, B.D., 2015].

Source Literature:

Adams, A. & Reeve, L. A. (1850). The zoology of the voyage of H.M.S. Samarang, under the command of Captain Sir Edward Belcher, C.B., F.R.A.S., F.G.S., during the years 1843-1846. Reeve & Benham, London. https://www.biodiversitylibrary.org/page/39770936 [Accessed 1 April 2025]
Brown, L. & Neville, B.D. (2015). Catalog of the recent taxa of the families Epitoniidae and Nystiellidae (Mollusca: Gastropoda) with a bibliography of the descriptive and systematic literature. Zootaxa. 3907(1). Magnolia Press Auckland, New Zealand. http://dx.doi.org/10.11646/zootaxa.3907.1.1 [Accessed 22 October 2023]
Clessin, S. (1897). Die Familie der Scalariidae. In W. Kobelt (Ed.), Systematisches Conchylien-Cabinet von Martini und Chemnitz. Neu herausgegeben und vervollständigt. 2(13). Bauer & Raspe, NürnbergBauer & Raspe, Nürnberg. https://www.biodiversitylibrary.org/page/34259905 [Accessed 21 June 2025]
Higo, S., Callomon, P. & Goto, Y. (1999). Catalogue and Bibliography of the Marine Shell-Bearing Mollusca of Japan.. https://www.researchgate.net/publication/266375656_Catalogue_and_Bibliography_of_the_Marine_Shell-Bearing_Mollusca_of_Japan [Accessed 25 July 2025]
Kaicher, S.D. (1980). Epitoniidae I [Pack 23]. S. D. Kaicher, St. Petersburg, Florida. http://www.femorale.com/kaicher/species.asp?f=Epitoniidae&c=125 [Accessed 8 May 2023]
NMW National Museum Wales (n.d.). Scalaria inclyta Melvill, 1885 Holotype. Amgueddfa Cymru - National Museum Wales, Cardiff. https://gbmolluscatypes.ac.uk/specimens/974 [Accessed 30 March 2025]
Sowerby, G. B. II. (1873-1874). Monograph of the genus Scalaria. Conchologia iconica, or illustrations of the shells of molluscous animals, vol. 19, pls 1-16 and unpaginated text.. 19. London. https://www.biodiversitylibrary.org/page/8217819 [Accessed 24 August 2023]
Tryon, G.W. (1887). Manual of conchology, structural and systematic with illustrations of the species. IX. Academy of Natural Sciences. https://www.biodiversitylibrary.org/page/11026889 [Accessed 20 September 2025]
Wilson, B.R. (1993). Australian Marine Shells. Australian Marine Shells. 1. Odessey Publications
WoRMS Editorial Board (2025). World Register of Marine Species. https://www.marinespecies.org [Accessed 8 November 2025]

WoRMS direct page link: Open WoRMS record

Epitonium pallasi neglectum

(A. Adams & Reeve, 1850)

Description:

Distribution:

Japan, south to the Philippines and Queensland, Australia

Habitat:

50-230m, sand

Etymology:

[latin] from neglectus = having been neglected, ignored, or overlooked

Type Material:

Distribution

Adams, A. & Reeve, L. A. (1850).

Plate VI. Fig. 5.

pp: 51.

Sowerby, G. B. II. (1873-1874).

Shown in text as Scalaria neglecta A. Adams & Reeve, 1850

Plate i. Fig. 1.

Fig. sp1.

Tryon, G.W. (1879).

Shown in text as Scalaria neglecta A. Adams & Reeve, 1850

Plate 11. Fig. 34.

Plate 11. Fig. 35.

pp: 55.

Tryon, G.W. (1879).

Shown in text as Scalaria inclyta Melvill, 1885

Plate 11. Fig. 35.

pp: 55.

Clessin, S. (1897).

Shown in text as Scalaria neglecta A. Adams & Reeve, 1850

Plate 7. Fig. 6.

pp: 27.

Shell broadly pyramidal, perforate, pale yellow between the varices; whorls increasing rapidly, disjointed; varices somewhat numerous, rather thin, continuous; near the suture drawn out into spines.

Kaicher, S.D. (1980).

Card #: EP1-2388.

Considered by some workers merely a variety (or synonym) of pallasi and, by others, a good species; it differs by having more and strongly hooked costae per whorl; to 40 Card #: EP1-2388

Wilson, B.R. (1993).

Shown in text as Epitonium neglectum (A. Adams & Reeve, 1850) ·

pp: 379. Fig. 31.

Nakayama, T. (2003).

pp: 123. Plate 10. Fig. 23.

h: 28.8 mm w: 15.0 mm

NMW National Museum Wales (n.d.).

Shown in text as Scalaria inclyta Melvill, 1885

Synonymy:

Source Literature:

WoRMS direct page link: Open WoRMS record

Considered by some workers merely a variety (or synonym) of pallasi and, by others, a good species; it differs by having more and strongly hooked costae per whorl; to 40

Card #: EP1-2388