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Epitoniidae Berry, 1910 (1812)

Variciscala raricostata

(G. B. Sowerby II, 1844)

Description:

Dimensions range from 18 mm - 25 mm high and 12 mm - 15.5 mm wide.
Width to height ratio: 0.67:1 ~ 0.62:1

Shell: pyramidal, ventricose; solid, thick; small to moderately small. Exterior colour white. About 5 whorls, rounded, rapidly increasing, convex, distinct. Suture deeply impressed, distinct. Umbilicate: narrow, small, but more or less obstructed by varices that extend inward. Approximately 45 costae on the body whorl. Costae are regularly spaced, low, subequal axial cords, with rare, prominent, crenulated, rounded, very few, unequally distant. rounded, crenulated, thick, thin, cord like. There is a single varix on each whorl. The varix is with an occasional thick, crenulated, reflected varix . Costae reflexed. Intercostal spaces numerous radial striae, creating a cancellate appearance where they intersect the costae. Fasciole complete. Aperture large, sub-circular, lip thickened by a weakly nodulose varix; inner lip slightly expanded anteriorly.


Distribution:

North Queensland, Australia [Wilson, 1993:281]; Sri Lanka, Mauritius [Brown & Neville, 2015:91]; Gulf of Oman [Melvill & Standen, 1903a:348]. Kiener [1873a:8] cites a specimen from Sicily; however, no corroborating evidence from other authors has been able to be found.


Habitat:

16.5-22 m [Iredale, 1930: 87], coral sand [Melvill & Standen, 1903a:348].


Etymology:

from rarus [latin] = scattered, far apart, seldom, rare or few and costatus = ribbed, in reference to the occasional expanded rib on the shell.


Type Material:

Brown & Neville [2015:91] regarded Cirsotrema kelea Iredale, 1930 as a junior synonym of Variciscala raricostata (G. B. Sowerby II, 1844). They also stated that Iredale's specimen was presumably held in the Australian Museum. However, a search of the Australian Museum's online catalogue confirmed the holotype of Cirsotrema kelea Iredale, 1930 as Australian Museum C.57741.

Holotype: Australian Museum: C.57741 [as Cirsotrema kelea Iredale, 1930] - Type Locality: Australia, Queensland, GBR, Michaelmas Cay (16° 36' 00' S, 145° 59' 00' E) /05/1926 - /06/1926, T.Iredale & G.P.Whitley(Collector), Great Barrier Reef Boring Expedition. [Australian Museum, 2026: C.57741]


SpeciesImage

Iredale, T. (1930). Plate X. Fig. 16.

Cirsotrema kelea Iredale, 1930

Distribution

as listed in source literature

Sowerby, G. B. II. (1844).
Species Image
Plate xxxiv. Fig. 79-80.
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pp: 93.
original description
Reeve, L.A. (1873).
Species Image
Plate ii. Fig. 9.
Species Image
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Plate xiv. Fig. 9b.
Kiener, L. C., & Fischer, P (1873a).
Shown in text as Scalaria raricosta Lamarck, 1822
Species Image
Plate II. Fig. 6.
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Plate II. Fig. 6.
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pp: 8.
6. SCALARIA CÔTES BARRÉES, Scalaria raricosta, LaM.
(Collection Mass. LAM.)
Sowerby, Genera, cahier 11, plate 2, fig. 3.
Plate II, fig. 6.
Description (Latin):
Shell turreted, perforated, white, with very fine transverse striae, obsolete longitudinal ribs; varices shaped like ribs, interrupted, rare and densely clustered in particular spots.
Description (French):
Shell oval-conical, somewhat turreted, cyclostome-shaped, with a pointed spire made up of six very convex, rounded whorls, on which some varices appear representing former peristomes; sometimes these varices are opposite one another, sometimes irregularly scattered. The whorls are separated by a rather deep suture. On these whorls, a very large number of small, closely spaced longitudinal striae can be seen, crossed by even finer transverse striae that give the shell a somewhat rough texture and form a rather regular fine mesh. The last whorl is subglobular, ending at its base in a small, narrow, and shallow umbilicus. The aperture is oval, rounded, almost semi-lunar. The peristome is thick, striated, and externally bordered by a convex ridge. The external color of the shell is a dull white.
Length: 9 lignes (about 20.3 mm or 0.8 inches).
Habitat: Mediterranean? Coasts of Sicily (Costa, Catalogue of Sicily).
Remarks:
This very peculiar shell is quite distinct from all other Scalaria species; in terms of external shape, it shows a strong resemblance to our elegant Cyclostome.
Tryon, G.W. (1887).
Shown in text as Scalaria raricostata Lamarck 1822, Scalaria reticulata Helbling, 1779
Species Image
Plate 16. Fig. 2-3.
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pp: 78.
Clessin, S. (1897).
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Plate 3. Fig. 6.
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pp: 10.
Shell white, thick, cancellated, pyramidal, swollen; whorls convex, distinct, rapidly increasing; varices very few, unevenly spaced, rounded, crenulated, thick; aperture rather large; inner lip slightly expanded at the front; umbilicus minute.
Iredale, T. (1930).
Shown in text as Cirsotrema kelea Iredale, 1930
Species Image
Plate X. Fig. 16.
Cirsotrema kelea Iredale, 1930
Species Image
pp: 87.
Mermod, G. (1963).
Shown in text as Scalaria raricosta Lamarck, 1822
Species Image
pp: 16+. Fig. 231.
Species Image
pp: 166.
Species Image
pp: 167.
Australian Museum (2026).
Shown in text as Cirsotrema kelea Iredale, 1930
Species Image
Fig. C.57741.
Cirsotrema kelea Iredale, 1930
Holotype: C.57741
Australia, Queensland, GBR, Michaelmas Cay (16° 36' 00" S, 145° 59' 00" E) /05/1926 - /06/1926, T.Iredale & G.P.Whitley(Collector), Great Barrier Reef Boring Expedition
Australian Museum. Photograph: M. Allen.
Copyright Australian Museum. Released under Creative Commons Licence CC BY-NC 4.0.

Synonymy:

Cirsotrema kelea Iredale, 1930 [Brown & Neville, 2015: 91; Australian Museum, 2026: C.57741; Iredale, 1930: 87]
Cirsotrema rariforme Abbott & Dance, 1982 [Brown & Neville, 2015: 129]
Scala raricosta Lamarck, 1822 [Melvill & Standen, 1903a: 348]
Scalaria raricosta Lamarck, 1822 [Lamarck, 1822: 228; Kiener & Fischer, 1873a: 8; Mermod & Binder, 1963: 166]
Scalaria raricostata Lamarck 1822 [Tryon, 1887: 78]
Scalaria raricostata Lamarck 1822 [Sowerby, 1844: 93; Clessin, 1897: 10; Reeve, 1873: pl. 2, fig. 9a-b]
Turbo raricostatus, "W. Wood", 1828 [Coan & Petit, 2011: 39]


Source Literature:

Australian Museum (2026). Australian Museum provider for OZCAM. Occurrence dataset. Atlas of Living Australia. https://doi.org/10.15468/e7susi accessed via GBIF.org [Accessed 18 July 2026]
Brown, L. & Neville, B.D. (2015). Catalog of the recent taxa of the families Epitoniidae and Nystiellidae (Mollusca: Gastropoda) with a bibliography of the descriptive and systematic literature. Zootaxa. 3907(1). Magnolia Press Auckland, New Zealand. http://dx.doi.org/10.11646/zootaxa.3907.1.1 [Accessed 22 October 2023]
Clessin, S. (1897). Die Familie der Scalariidae. In W. Kobelt (Ed.), Systematisches Conchylien-Cabinet von Martini und Chemnitz. Neu herausgegeben und vervollständigt. 2(13). Bauer & Raspe, NürnbergBauer & Raspe, Nürnberg. https://www.biodiversitylibrary.org/page/34259905 [Accessed 21 June 2025]
Coan, E.V. & Petit, R.E. (2011). The publications and malacological taxa of William Wood (1774-1857).. Malacologia. 54. http://dx.doi.org/10.4002/040.054.0109 [Accessed 25 October 2025]
Iredale, T. (1930). Queensland molluscan notes, No. 2.. Memoirs of the Queensland Museum.. 10(1). https://www.biodiversitylibrary.org/page/52128623 [Accessed 2 August 2026]
Kiener, L. C. & Fischer, P. (1873a). Spécies général et iconographie des coquilles vivantes comprenant la collection du Muséum d'histoire naturelle de Paris: la collection Lamarck, celle du prince Masséna (appartenant maintenant a M.B. Delessert) et les découvertes récentes des voyageurs. 10. J.B.Baillière et Fils. https://www.biodiversitylibrary.org/item/19879 [Accessed 19 April 2025]
Melvill, J. C. & Standen, R. (1903a). The genus Scala (Klein) Humphrey, as represented in the Persian Gulf, Gulf of Oman, and North Arabian Sea, with descriptions of new species.. Journal of Conchology. 10. Conchological Society of Great Britain and Ireland. https://www.biodiversitylibrary.org/page/31650162 [Accessed 26 February 2025]
Mermod, G. & Binder, E. (1963). Les types de la Collection Lamarck au Muséum de Genève. (Mollusques vivants, V). Revue suisse de zoologie. 70(7). https://doi.org/10.5962/bhl.part.75362 [Accessed 4 August 2025]
Reeve, L. A. (1873). Monograph of the genus Scalaria. Conchologia iconica, or illustrations of the shells of molluscous animal. 19. London. https://www.biodiversitylibrary.org/page/8217819 [Accessed 24 August 2023]
Sowerby, G. B. II. (1844). Monograph of the genus Scalaria.. Thesaurus conchyliorum, or monographs of genera of shells.. 1 (4). London, privately published.. http://biodiversitylibrary.org/page/11076419 [Accessed 22 June 2023]
Tryon, G.W. (1887). Manual of conchology, structural and systematic with illustrations of the species. IX. Academy of Natural Sciences. https://www.biodiversitylibrary.org/page/11026889 [Accessed 20 September 2025]
Wilson, B.R. (1993). Australian Marine Shells. Australian Marine Shells. Vol.1. Odessey Publications
WoRMS Editorial Board (2025). World Register of Marine Species. https://www.marinespecies.org at VLIZ. [Accessed 8 November 2025]

WoRMS direct page link: Open WoRMS record