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Epitoniidae Berry, 1910 (1812)

Filiscala raricosta

(Lamarck, 1804)

Description:

Dimensions range from 5.4 mm - 25 mm high and 5.98 mm - 7.66 mm wide.
Width to height ratio: 01.11:1 ~ 0.31:1

Shell: pyramidal, pointed; thin; minute to moderately small. Exterior colour white, costae white. About 12 whorls, convex, rounded whorls that bulge outward significantly, with the most prominent outer edge sitting around the middle; protoconch with ~3 whorls ; teleoconch with ~7 whorls. Suture impressed. Umbilicate: narrow, partially obscured by peristome. Approximately 17 - 25 costae on the body whorl. Costae are very fine or obsolete varices, some thick, and curved strong,, varices very few, rounded, crenated, slightly curved above the middle;, threadlike, irregularly spaced, low rounded ribs. There is a single varix on each whorl. The varix is with occasional thicker varices. Costae aligned on earlier whorls, but discontinuous on later whorls; discontinuous across the suture; not peaked; not reflexed. Intercostal spaces fine, evenly spaced radial threads. Interstices with approximately 20 spiral striae per whorl. Base with rounded striae. Aperture oval, ear-shaped, large; peristome thickened and reflected but incomplete; inner lip labium moderately expanded over umbilical opening.

Tryon [1887: 78] notes that Reeve's locality, West Indies, is erroneous.


Distribution:

Indo-West Pacific, India to Gulf of Oman [Ghosh et al, 2016:482]; Cocoss Keeling Islands [Maes, 1967:117]; Singapre [Tan & Woo, 2010:40]; Australia, Queensland south to Keppel Bay: [Wilson, 1990: 279]; Wexmouth , Western Australia [South Australian Museum, 2026: D51462]; Japan, southward from Kii Peninsular [Higo et al.,1999]; Taiwan [Huang & Lee, 2016:19]; Philippines, Mauritius [Tryon, 1887:78].


Habitat:

offshore, 20 [Higo et al., 1999] to 285.3m [Melvill & Standen, 1903a:348]; sandy mud [Higo et al., 1999].


Etymology:

[latin] from rarus = scattered, far apart, seldom, rare or few and costatus = ribbed, in reference to the occasional expanded rib on the shell.


SpeciesImage

Ghosh, A., Mukhopadhay, A. & Tripathy, B (2016). pp: 482. Fig. 2.

Fig. 2. Filiscala raricosta (Lamarck, 1804)

Distribution

as listed in source literature

Type Material:

The following list brings together type material and other cited specimens as reported in the sources examined, grouped under the names and designations used in each source. The same specimen may appear in multiple entries drawn from different sources. Some entries concern other species included by authors for comparison; inclusion in this list does not imply synonymy.
Filiscala martinii (W. Wood, 1828)Type designation not provided in text/ source documentnone designated. [Higo et al., 1999: 223].
Scalaria striata Kiener, 1838Type not locatedno location given, [Brown & Neville, 2015: 142].
Turbo martinii W. Wood, 1828HolotypeNHM 20100591. [Natural History Museum, 2014: NHM 20100591].NHMUK 20100591. [Coan & Petit, 2011: 39].TypeNHMUK 20100591. no location given, [Brown & Neville, 2015: 102].


Chemnitz, J. H. (1780).
Species Image
Plate 153. Fig. 1435.
Species Image
Plate 153. Fig. 1436.
Lamarck [J.B.P.A. de]. (1804).
Species Image
pp: 213.
All Scalaria are marine shells. Some inhabit the seas of warm climates, while others occur in the seas bordering our oceanic coasts.
Among the species of this genus, one is particularly remarkable for its umbilicus and for the unusual separation of the whorls of its spire. It is highly sought after by collectors and, for this reason, commands a fairly high price in trade, especially when it is of large size and well preserved.
This is Linnaeus's Turbo scalaris, the species to which the name scalata is more especially applied. To this genus should also be referred Turbo principalis Pallas, Turbo clathrus Linnaeus
Lamarck, [J.-B. M.] de. (1822).
Shown in text as Scalaria raricosta.
Species Image
pp: 228.
Kiener, L. C., & Fischer, P. (1838b).
Shown in text as Scalaria striata Kiener, 1838
Species Image
Plate V. Fig. 14.
Species Image
Plate V. Fig. 14.
Species Image
pp: 20.
Sowerby, G. B. II. (1844).
Shown in text as Scalaria martinii (W. Wood, 1828)
Species Image
Plate iii. Fig. 20.
Species Image
Reeve, L.A. (1873).
Shown in text as Scalaria martinii (W. Wood, 1828)
Species Image
Plate xxxiv. Fig. 78.
Species Image
pp: 93.
Tryon [1887: 78] notes that Reeve's locality, West Indies, is erroneous.
Tryon, G.W. (1887).
Shown in text as Scalaria martinii Wood, 1828
Species Image
Plate 16. Fig. 5.
Species Image
pp: 78.
Clessin, S. (1897).
Shown in text as Scalaria martinii (W. Wood, 1828)
Species Image
Plate 8. Fig. 9.
Species Image
pp: 31.
Shell elevated, white, spirally striated; whorls numerous, rounded, contiguous; varices very few, rounded, crenulated, slightly arched above the middle; aperture somewhat auriform.
Melvill, J. C. & Standen, R. (1903a).
Shown in text as Scala (Amaea) martinii (W. Wood, 1828)
Species Image
pp: 348.
Maes, V. O. (1967).
Shown in text as Epitonium martinii (W. Wood, 1828)
Species Image
Plate 8. Fig. c.
Epitonium martinji (Wood)
ANSP 287713, 5.4 mm
Coan, E.V. & Petit, R.E. (2011).
Shown in text as Turbo martinii, W. Wood, 1828
Species Image
pp: 39.
Ghosh, A., Mukhopadhay, A. & Tripathy, B (2016).
Species Image
pp: 482. Fig. 2.
Fig. 2. Filiscala raricosta (Lamarck, 1804)

Synonymy:

Amaea (Filiscala) martinii (W. Wood, 1828) [Weil et al., 1999: 110]
Amaea martinii (W. Wood, 1828) [Kaicher, 1980: EP1-2335]
Epitonium martinii (W. Wood, 1828) [Maes, 1967: nd]
Filiscala martinii (W. Wood, 1828) [Wilson, 1993: 379; Higo et al., 1999: 223; Tan & Woo, 2010: 40]
Scala (Amaea) martinii (W. Wood, 1828) [Melvill & Standen, 1903a: 348]
Scalaria martinii (W. Wood, 1828) [Sowerby, 1844: 93; Sowerby, 1844: ND; Clessin, 1897: 31]
Scalaria martinii Wood, 1828 [Tryon, 1887: 78]
Scalaria raricosta Lamarck, 1822 [Lamarck, 1822: 228; Kiener & Fischer, 1873a: 8; Mermod & Binder, 1963: 166]
Scalaria striata Kiener, 1838 [Brown & Neville, 2015: 142; Kiener & Fischer, 1838b: 20]
Turbo martinii W. Wood, 1828 [Brown & Neville, 2015: 102; Coan & Petit, 2011: 39; Natural History Museum, 2014: NHM 20100591]


Source Literature:

arsint.com (2023). Shells for Sale:Epitonidae. arsint.com. http://www.arsint.com/seashells_epitoniidae_opalia.html [Accessed 21 October 2023]
Brown, L. & Neville, B.D. (2015). Catalog of the recent taxa of the families Epitoniidae and Nystiellidae (Mollusca: Gastropoda) with a bibliography of the descriptive and systematic literature. Zootaxa. 3907(1). Magnolia Press Auckland, New Zealand. http://dx.doi.org/10.11646/zootaxa.3907.1.1 [Accessed 22 October 2023]
Chemnitz, J. H. (1780). Neues systematisches Conchylien-Cabinet. Neues systematisches Conchylien-Cabinet. 4. Nürnberg: Gabriel Nicolaus Raspe. https://upload.wikimedia.org/wikipedia/commons/1/19/Neues_systematisches_Conchylien-Cabinet._Bd._04%2C_1780_%28IA_b30414994_0004%29.pdf [Accessed 20 March 2025]
Clessin, S. (1897). Die Familie der Scalariidae. In W. Kobelt (Ed.), Systematisches Conchylien-Cabinet von Martini und Chemnitz. Neu herausgegeben und vervollständigt. 2(13). Bauer & Raspe, NürnbergBauer & Raspe, Nürnberg. https://www.biodiversitylibrary.org/page/34259905 [Accessed 21 June 2025]
Coan, E.V. & Petit, R.E. (2011). The publications and malacological taxa of William Wood (1774-1857).. Malacologia. 54. http://dx.doi.org/10.4002/040.054.0109 [Accessed 25 October 2025]
Ghosh, A., Mukhopadhay, A. & Tripathy, B (2016). New Records of Two Epitoniids Cycloscala crenulata (Pease, 1896), Filiscala raricosta (Lamarck, 1804) (Gastropoda: EPITONIIDAE) From the West Coast of India. Records of the Zoological Survey of India. 116(4). https://www.researchgate.net/publication/344238483_Short_Communication_NEW_RECORDS_OF_TWO_EPITONIIDS_CYCLOSCALA_CRENULATA_PEASE_1896_FILISCALA_RARICOSTA_LAMARK_1804_GASTROPODA_EPITONIIDAE_FROM_THE_WEST_COAST_OF_INDIA [Accessed 16 September 2023]
Higo, S., Callomon, P. & Goto, Y. (1999). Catalogue and Bibliography of the Marine Shell-Bearing Mollusca of Japan.. https://www.researchgate.net/publication/266375656_Catalogue_and_Bibliography_of_the_Marine_Shell-Bearing_Mollusca_of_Japan [Accessed 25 July 2025]
Huang, C. W. & Lee, Y.C. (2016). Checklist of the family Epitoniidae (Mollusca: Gastropoda) in Taiwan with description of a new species and some new records. Biodiversity Data Journal. 4. doi: 10.3897/ BDJ.4.e5653 [Accessed 25 August 2023]
Kaicher, S.D. (1980). Epitoniidae I [Pack 23]. S. D. Kaicher, St. Petersburg, Florida. http://www.femorale.com/kaicher/species.asp?f=Epitoniidae&c=125 [Accessed 8 May 2023]
Kiener, L. C. & Fischer, P. (1838b). Genre Scalaire. Spécies général et iconographie des coquilles vivantes comprenant la collection du Muséum d’histoire naturelle de Paris : la collection Lamarck, celle du prince Masséna (appartenant maintenant a M.B. Delessert) et les découvertes récentes des voyageurs. 10. J.B.Baillière et Fils. http://www.biodiversitylibrary.org/item/19879 [Accessed 11 May 2026]
Lamarck [J.B.P.A. de]. (1804). Suite des mémoires sur les fossiles des environs de Paris.. Annales du Muséum d'Histoire Naturelle. 4. https://biodiversitylibrary.org/page/29642493 [Accessed 30 July 2026]
Lamarck, [J.-B. M.] de. (1822). Histoire naturelle des animaux sans vertèbres.. Histoire naturelle des animaux sans vertèbres.. 6(2). Lamarck, [J.-B. M.] de.. http://biodiversitylibrary.org/page/13181853 [Accessed 20 March 2025]
Maes, V. O. (1967). The Littoral Marine Mollusks of Cocos-Keeling Islands (Indian Ocean). Proceedings of the Academy of Natural Sciences of Philadelphia. 119. Academy of Natural Sciences. https://www.jstor.org/stable/4064613 [Accessed 25 October 2025]
Melvill, J. C. & Standen, R. (1903a). The genus Scala (Klein) Humphrey, as represented in the Persian Gulf, Gulf of Oman, and North Arabian Sea, with descriptions of new species.. Journal of Conchology. 10. Conchological Society of Great Britain and Ireland. https://www.biodiversitylibrary.org/page/31650162 [Accessed 26 February 2025]
Murrell, T. (2023). Personal Collection
Natural History Museum (2014). NHM Collection specimens [Data set]. Natural History Museum. https://doi.org/10.5519/0002965 [Accessed 6 April 2026]
South Australian Museum (2026). South Australian Museum Adelaide provider for OZCAM. Atlas of Living Australia. https://doi.org/10.15468/wz4rrh accessed via GBIF.org [Accessed 21 July 2026]
Sowerby, G. B. II. (1844). Monograph of the genus Scalaria.. Thesaurus conchyliorum, or monographs of genera of shells.. 1 (4). London, privately published.. http://biodiversitylibrary.org/page/11076419 [Accessed 22 June 2023]
Tan, S. K. & Woo, H. P. M. (2010). A preliminary checklist of the molluscs of Singapore. Raffles Museum of Biodiversity Research & Department of Biological Sciences, National University of Singapore.
Tryon, G.W. (1887). Manual of conchology, structural and systematic with illustrations of the species. IX. Academy of Natural Sciences. https://www.biodiversitylibrary.org/page/11026889 [Accessed 20 September 2025]
Weil, A., Brown, L. & Neville, B. (1999). The Wentletrap Book. Evolver
Wilson, B.R. (1993). Australian Marine Shells. Australian Marine Shells. Vol.1. Odessey Publications
WoRMS Editorial Board (2025). World Register of Marine Species. https://www.marinespecies.org at VLIZ. [Accessed 8 November 2025]

WoRMS direct page link: Open WoRMS record