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Epitoniidae Berry, 1910 (1812)

Epitonium lyra

(Sowerby II, 1844)

Description:

Dimensions range from 6.7 mm - 94.8 mm high and 3.38 mm - 15 mm wide.
Width to height ratio: 0.5:1 ~ 0.16:1

Shell: broadly pyramidal, stout; thin, fragile; minute to large. Exterior colour very pale brown to dull white, with two bands of deeper or paler brown, costae white. About 9 whorls, inflated, rounded, connected; teleoconch with ~6 whorls relatively low spire, inflated. Spire relatively low, convex. Suture deep. Umbilicate: narrow, moderate. Approximately 37 - 45 costae on the body whorl. Costae are prosocline, low, thin and sharp, curving into the umbilicus. Costae not alinged; not peaked; reflexed crests; not continuous across the suture. Intercostal spaces wth minute spiral striae. Aperture oblong-ovate, ear-shaped; peristome complete, thin, simple.


Distribution:

Indo-West Pacific; Amani O-shima to Okinawa, Japan; East China Sea, Philippines, Papua New Guinea, Fiji; Port Douglas to Capricorn Bunker Group Queensland, Australia; Red Sea, Mozambique


Habitat:

intertidal to 522m (Schepman 1909: 224), sand, stony substrate


Etymology:

from lyra [latin] = lyre or lute, in possible reference to the shell's costae resembling the strings of a lyre


Type Material:

Syntype: (BM(NH) 1979162) for Epitonium lyra (Sowerby II, 1844) - Type Locality: from Isle of Masbate, Philippines, on the reefs; the only undamaged shell is here designated lectotype.. [Kilburn, R.N, 1985]
Lectotype: NHMUK 1979162/1-3 (lectotype, paralectotypes) selected by Kilburn (1985: 309). for Epitonium lyra (Sowerby II, 1844) - Type Locality: Masbate Island, Philippines.. [Brown, L. & Neville, B.D., 2015]
Type not located. [as Scalaria vestalis Hinds, 1844:] - Type Locality: New Guinea, seven fms [=13 m], among mud.. [Brown, L. & Neville, B.D., 2015]
Type not listed in source data for Epitonium lyra (Sowerby II, 1844) - Type Locality: China Sea. [Higo, S., Callomon, P. & Goto, Y., 1999]


SpeciesImage

Sowerby, G. B. II. (1844). pp: xxxii. Plate xxxiii. Fig. 38-39.

Distribution

as listed in source literature

Sowerby, G. B. II. (1873).
Shown in text as Scalara vestalis
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Plate xxxiv. Fig. 97.
was Sc.vestalis
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pp: 93.
38. Sc. vestalis. (pi. xxxiv f. 97.) Hinds. Zool. Proc. 1843.
Sowerby, G. B. II. (1844).
Shown in text as Scalaria lyra
Species Image
pp: xxxii. Plate xxxiii. Fig. 38-39.
Species Image
pp: 98.
original description
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pp: 90.
original description
Sowerby, G. B. II. (1873).
Shown in text as SCALARIA VESTALIS.
Species Image
Plate xiii. Fig. 101.
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Sowerby, G. B. II. (1873).
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Plate iv. Fig. 23.
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Species 23. (Mus. Brit.)
SCALARIA LYRA.
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Tryon, G.W. (1879).
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Plate 12. Fig. 69.
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pp: 58.
Tryon, G.W. (1887).
Shown in text as Scalaria vestalis Hinds, 1844
Species Image
Plate 14. Fig. 37.
Species Image
Clessin, S. (1897).
Species Image
Plate 6. Fig. 4.
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pp: 22.
ventricose, rather short, pale tawny, with two tawny spiral bands, with extremely narrow, lamellar varices; very densely ridged; whorls rounded, contiguous; aperture ear-shaped.
Clessin, S. (1897).
Shown in text as Scalaria vestalis Hindson
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Plate 13. Fig. 8.
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pp: 52.
Shell small, whitish with a bluish tinge, shortly pyramidal; spirally very finely striated; whorls rounded; varices simple, minute; aperture large, ovate, rather broad.
Schepman, M.M. (1909).
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pp: 224.
Kaicher, S.D. (1980).
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Card #: EP1-2341.
USNM: 343441
Kilburn, R.N (1985).
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122, Lectotype, BM(NH) 1979162
21,2 x 13 mm;
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123, NM J5476,
Benguera Is., 14,5 x 9,1 mm.
Wilson, B.R. (1993).
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pp: 379. Plate 44. Fig. 28.
Nakayama, T. (2003).
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pp: 139. Plate 18. Fig. 13.
Epitonium (Limiscala) lyra (Sowerby, 1844)
height 22.0 mm; breadth 15.0 mm (WMNH-Mo-Na676)

Synonymy:

Scala lyra G. B. Sowerby II, 1844 [in Thiele, J., 1925].
Scalaria lyra [in Sowerby, G. B. II., 1844].
Scalaria vestalis Hinds, 1844 [in Hinds, R. B., 1843].
Scalaria vestalis Hinds, 1844 [in Sowerby, G. B. II., 1844].
Scalaria vestalis Hinds, 1844 [in Sowerby, G. B. II., 1873].
Scalaria vestalis Hinds, 1844 [in Tryon, G.W., 1887].
Scalaria vestalis Hinds, 1844: [in Brown, L. & Neville, B.D., 2015].
Scalaria vestalis Hindson [in Clessin, S., 1897].


Source Literature:

Brown, L. & Neville, B.D. (2015). Catalog of the recent taxa of the families Epitoniidae and Nystiellidae (Mollusca: Gastropoda) with a bibliography of the descriptive and systematic literature. Zootaxa. 3907(1). Magnolia Press Auckland, New Zealand. http://dx.doi.org/10.11646/zootaxa.3907.1.1 [Accessed 22 October 2023]
Clessin, S. (1897). Die Familie der Scalariidae. In W. Kobelt (Ed.), Systematisches Conchylien-Cabinet von Martini und Chemnitz. Neu herausgegeben und vervollständigt. 2(13). Bauer & Raspe, NürnbergBauer & Raspe, Nürnberg. https://www.biodiversitylibrary.org/page/34259905 [Accessed 21 June 2025]
Higo, S., Callomon, P. & Goto, Y. (1999). Catalogue and Bibliography of the Marine Shell-Bearing Mollusca of Japan.. https://www.researchgate.net/publication/266375656_Catalogue_and_Bibliography_of_the_Marine_Shell-Bearing_Mollusca_of_Japan [Accessed 25 July 2025]
Hinds, R. B. (1843). Descriptions of new species of Scalaria and Murex, from the collection of Sir Edward Belcher, C.B.. Proceedings of the Zoological Society of London.. 11. http://biodiversitylibrary.org/page/30680102 [Accessed 28 December 2025]
Kaicher, S.D. (1980). Epitoniidae I [Pack 23]. S. D. Kaicher, St. Petersburg, Florida. http://www.femorale.com/kaicher/species.asp?f=Epitoniidae&c=125 [Accessed 8 May 2023]
Kilburn, R.N (1985). The family Epitoniidae (Mollusca: Gastropoda) in southern Africa and Mozambique. Annals of the Natal Museum. 27(1). Natal Museum
Nakayama, T. (2003). A Review of Northwest Pacific Epitoniids. Monographs of Marine Mollusca. 6. Backhuys Publishers
Schepman, M.M. (1909). The Prosobranchia of the Siboga Expedition. Part 2. Taenioglossa and Ptenoglossa. E.J. Brill,Leyden. https://www.biodiversitylibrary.org/page/34314807 [Accessed 27 July 2025]
Sowerby, G. B. II. (1844). Monograph of the genus Scalaria.. Thesaurus conchyliorum, or monographs of genera of shells.. 1 (4). London, privately published.. http://biodiversitylibrary.org/page/11076419 [Accessed 22 June 2023]
Sowerby, G. B. II. (1873). Monograph of the genus Scalaria. Conchologia iconica, or illustrations of the shells of molluscous animals, vol. 19, pls 1-16 and unpaginated text.. 19. London. https://www.biodiversitylibrary.org/page/8217819 [Accessed 24 August 2023]
Thiele, J. (1925). Gastropoden der Deutschen Tiefsee-Expedition. II Teil. Wissenschaftliche Ergebnisse der Deutschen Tiefsee-Expedition auf dem Dampfer "Valdivia" 1898-1899.. 17(2). https://www.deutsche-digitale-bibliothek.de/item/3NJJP6TFO3ZWXZDRTI2CZV66VP76UDWR [Accessed 26 October 2025]
Tryon, G.W. (1887). Manual of conchology, structural and systematic with illustrations of the species. IX. Academy of Natural Sciences. https://www.biodiversitylibrary.org/page/11026889 [Accessed 20 September 2025]
Weil, A., Brown, L. & Neville, B. (1999). The Wentletrap Book. Evolver
Wilson, B.R. (1993). Australian Marine Shells. Australian Marine Shells. 1. Odessey Publications
WoRMS Editorial Board (2025). World Register of Marine Species. https://www.marinespecies.org [Accessed 8 November 2025]

WoRMS direct page link: Open WoRMS record